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l஌ཱྀ֮ߦzͷͨΊͷࢹ֮୅ߦσόΠε ࿨Ոঘرླ໦ྑฏ ෬ݟྒྷฏफ૾༔ཬ "VEJUPSZWJTJPOTVCTUJUVUJPOEFWJDF GPSQFSDFQUJPOBMWPZBHF

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Ϟνϕʔγϣϯ ԿͷͨΊʹ4JHIUΛ ࡞ͬͨͷ͔

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νʔϜϝϯόʔ ླ໦ྑฏ फ૾༔ཬ ࿨Ոঘر ෬ݟྒྷฏ ౦ژେֶ৘ใཧ޻. ઐ໳6* ଟຎඒज़େֶ৘ใσβΠϯ# ઐ໳σβΠϯ ౦ژେֶ৘ใཧ޻. ઐ໳ਆܦՊֶ ౦ژେֶֶࡍ৘ใ. ઐ໳73ɾ৺ཧֶ

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νʔϜϝϯόʔ ླ໦ྑฏ फ૾༔ཬ ࿨Ոঘر ෬ݟྒྷฏ ౦ژେֶ৘ใཧ޻. ઐ໳6*ɾԻڹ ଟຎඒज़େֶ৘ใσβΠϯ# ઐ໳σβΠϯ ౦ژେֶ৘ใཧ޻. ઐ໳ਆܦՊֶ ౦ژେֶֶࡍ৘ใ. ઐ໳73ɾ৺ཧֶ ஌֮ ώτ͕ͲͷΑ͏ͳ࢓૊ΈͰɺͲͷΑ͏ʹ ΋ͷΛݟͯԻΛฉ͍͍ͯΔͷ͔

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஌֮ͷՄ઼ੑ Մ઼ੑೖྗͷมԽʹରͯ͠ ॊೈʹػೳΛൃش͢Δੑ࣭

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൓ڹԻʹΑΔԻڹఆҐ

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ٯ͞ϝΨωͷ࣮ݧ 4USBUUPO`T*OWFSUJOH(MBTTFT

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[Norimoto & Ikegaya, 2015] Ϛ΢εҰ࣍ࢹ֮໺ʹ࣓ؾ৘ใΛೖྗ͢Δͱ
 ޙఱతʹz࣓৔Λݟͯzۭؒ୳ࡧͰ͖ΔΑ͏ʹͳΔ

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10 ʹ ஌֮ͷՄ઼ੑ ೴ͷॊೈੑ

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→

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l஌ཱྀ֮ߦzͷͨΊͷࢹ֮୅ߦσόΠε "VEJUPSZWJTJPOTVCTUJUVUJPOEFWJDF GPSQFSDFQUJPOBMWPZBHF

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࣮૷ͷ֓ཁͱܦҢ ԻʹΑΔzݟ͑zͷମݧʜ ͲͷΑ͏ʹ࣮ݱ͢Δ͔

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΋ͬͱ΋ॏཁͳ՝୊ ֎քͷ৘ใͷͲ͜ͷ෦෼Λ Ͳ͏΍ͬͯԻʹม׵͢Δ͔ ΅ͨͪ͘͸ݟΔ͜ͱʹΑͬͯ ֎քʹؔ͢ΔͲΜͳ৘ใΛಘ͍ͯΔͷ͔

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͜Ε·Ͱͷ࣮૷ WFS WFS WFS WFS ௿࣍ͷಛ௃఺ɾہॴಛ௃ྔ 463' ͷ෼෍ த৺ը૾ΛҰൠ෺ମೝࣝ%//ʹೖྗͨ͠தؒ৘ใ ࢹ໺಺਺఺ͷਂ͞৘ใ ฏ໘഑ஔ͔ΒߦҝՄೳੑΛղੳ ը૾ͷ໨ཱͭྖҬΛࡉ͔۠͘੾ͬͯԻݯͱରԠͤ͞Δ ࣅ͍ͯΔ΋ͷ͕ɺࣅͨԻʹม׵͞ΕΔΑ͏ͳԻੜ੒ ۭؒߏ଄ʹண໨ɺσϓεΧϝϥΛಋೖ ۭؒߏ଄ͷҙຯతͳଆ໘ʹண໨͠ɺฏ໘நग़

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WFS w ը૾ͷಛʹ໨ཱͭ෦෼ಛ௃఺Λղੳ͠ɺ
 ͦͷ఺ͷपғͷಛ௃ΛɺԻݯʹରԠͤ͞Δ ಛ௃Λ Իʹม׵

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WFS ΦϨϯδ৭ͷ ෳࡶͳԻͷմ͕ খ͘͞ͳΓͳ͕Β ӈ͔ΒԞʹ޲͔͍ͬͯΔ Ͳ͏ฉ͑͜Δ͔?:

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WFSม׵ྫ τϯωϧ

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WFSߦಈ

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7FSҰൠ෺ମೝࣝͷ%//Λ༻͍ͨϞσϧ w ෺ମΛࣝผ͢ΔͨΊͷ%//ͷϞσϧͷ
 ࣝผͷख͕͔ΓͱͳΔதؒ૚ͷ৘ใΛநग़
 ˠԻੜ੒ͷύϥϝʔλͱͯ͠ར༻ w ࣅ͍ͯΔ΋ͷ͸ࣅ͍ͯΔԻʹ

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7FSࢹ໺಺਺఺ͷਂ͞৘ใ આ໌ प೾ w σϓεΧϝϥΛಋೖɺਂ͞৘ใΛ௚઀औಘ w ࢹ໺಺̑ํҐͦΕͧΕͷਂ౓෼෍Λύϥϝʔλʹ
 ϗϫΠτϊΠζΛมௐͯ͠Իڹ߹੒ w ԻͷมԽ͔Βো֐෺ͷҐஔ΍େ͖͕͞෼͔Δ

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7FSฏ໘഑ஔ͔ΒߦҝՄೳੑΛղੳ આ໌ น໘ʹҠಈͷ੍໿ ϘϦϡʔϜʹিಥɾ೺࣋ ฏ໘ʹ෺ମͷࢧ࣋ চ໘ʹҠಈൣғ w զʑ͕ݟ͍ͯΔͷ͸ۭؒͰ͸ͳ͘ߦҝͷՄೳੑ DG+(JCTPO w ۭؒΛ෼ੳɺ̐छྨͷجຊΦϒδΣΫτΛநग़ w ΦϒδΣΫτ͝ͱʹԻݯΛ഑ஔ

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7FSฏ໘഑ஔ͔ΒߦҝՄೳੑΛղੳ ࣮૷ ϙΠϯτΫϥ΢υͱͯ͠
 ฏ໘நग़ɾΫϥελϦ ϯά ΦϒδΣΫτ΁ͷԻݯ഑ஔ
 ཱମԻڹ৞ΈࠐΈ *3$". TQBU

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ϙΠϯτΫϥ΢υͷղੳྫ ϘϦϡʔϜ ਨ௚໘

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࣮ࡍͷ߹੒Իڹͷྫ

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Ծ૝؀ڥΛ༻͍ͨ4JHIUͷ࿅श w %্ۭؒͰɺ͋Δۭ͕ؒͲ͏ௌ͑͜Δͷ͔܇࿅

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ϋʔυ΢ΣΞͷมભ

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ϋʔυ΢ΣΞͷ੍໿ w %ϓϦϯλ "#4 ੡ w 9UJPO1SPηϯαΛ෇͚֎͠Մೳ w ϔουϗϯ༻υϥΠόɾΠϠύου಺ଁ w ͋ͱ͔Β૊Έཱͯɾ෼ղՄೳ w લޙࠨӈʹ಄Λৼͬͯ΋0,

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૊ΈཱͯػߏΛಋೖ

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࠷৽൛ w Մಈੑɺ଱ٱੑɺ҆ఆੑΛ޲্ w δϣΠϯτ෦෼ͷڧ౓ΛΞοϓ


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ͲΜͳ΋ͷΛ࡞ͬͨͷ͔ ධՁͱ஌ݟ

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λεΫιϑΝʹ࠲Δ

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λεΫนΛආ͚ͯา͘ ؟ΛݟுΔ΄Ͳ ૉ੖Β͍͠ σϞө૾

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λεΫ֊ஈΛͷ΅Δ ؟ΛݟுΔ΄Ͳ ૉ੖Β͍͠ σϞө૾

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λεΫਓΛ௥͍͔͚Δ ؟ΛݟுΔ΄Ͳ ૉ੖Β͍͠ σϞө૾

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λεΫΩϟονϘʔϧ ؟ΛݟுΔ΄Ͳ ૉ੖Β͍͠ σϞө૾

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ຊ౰ʹ͜Μͳ͜ͱ͕Ͱ͖Δͷ͔ 2

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ϥΠϒσϞ ਓͷ͍ΔҐஔɾڑ཭Λ೺Ѳ͠ ૬खʹ޲͔ͬͯϘʔϧΛ౤͛Δ

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௕࣌ؒண༻ͷ༷ࢠ

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4JHIUΛ௕࣌ؒண༻ͨ͠ͱ͖ͷมԽ ໨Ͱݟ͍ͯΔͱ͖ͷΑ͏ʹɺ ෺ମ͕ʮݎͯ҆͘ఆ࣮ͨ͠ମʯͰ͋Δ Α͏ʹ͸ײ͡ΒΕͳ͘ͳΔ l Ի͕ྲྀΕग़͢ݯͱͯ͠෺ମͷଘࡏΛ ײ͡ΒΕΔΑ͏ʹͳΔ z

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4JHIUΛ௕࣌ؒண༻ͨ͠ͱ͖ͷมԽ Կ͔ͷཧ༝ͰԻ͕ฉ͑͜ͳ͘ͳΔͱɺ ײ֮ث׭ΛୣΘΕͨΑ͏ʹ ͱͯ΋ෆ҆ͳؾ෼ʹͳΔ l z

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΄Μͱ͏ʹͩΕͰ΋ମݧͰ͖Δͷ͔ 2

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46 7FS ౦ژେ੍ֶ࡞ల

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47 7FS %$&910

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ϢʔβελσΟ w %$&910΍౦େɾ౦ژܳେͰͷలࣔΛ௨ͯ͠
 ͷ΂໊Ҏ্ͷํ͕4JHIUΛମݧ w ୹࣌ؒͷମݧͳ͕Βɺ্ୡͷ଎͍ਓ͸น΍صɺ
 ਓͷҐஔΛ೺ѲͰ͖ΔΑ͏ʹ w ΞϯέʔτͰଟ͔ͬͨײ૝ w ʮصͷน΍Ґஔ͕Θ͔ͬͨʯʮԻ͔Β·ΘΓͷޫܠΛ૝૾ͨ͠ʯ w ʮυΩυΩͨ͠ɾָ͔ͬͨ͠ʯʮί΢ϞϦʹͳͬͨΈ͍ͨͩͬͨʯ w ʮͲ͏͍͏࢓૊Έͳͷ͔ؾʹͳͬͨʯ

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ࢹ֮৘ใʹཔΒͣʹੜ׆Λ͍ͯ͠Δํ͸ɼ 4JHIUͷੈքΛͲ͏ײ͡Δ 2

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ϢʔβελσΟ w ࢁԼ͞Μ ࢹ֮ো֐౰ࣄऀ ઌఱੑશ໡ ࣸਅ

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ͲͷΑ͏ͳ൓ڹɾӨڹ͕͔͋ͬͨ ࠓޙͲͷΑ͏ͳల๬Λ͍࣋ͬͯΔͷ͔ ࣾձ΁ͷΠϯύΫτ

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ࣾձ΁ͷΠϯύΫτ ϝσΟΞܝࡌˠ໰͍߹Θͤˠ ϢʔβελσΟ΍࣍ͷలࣔʹܨ͕Δ

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ۮવͰ͕͢ l%..NBLFz࣌ؒີணऔࡐͰͷ์ө͔Β ϓϩμΫτ΁ͷҙݟ͕ू·Δ

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౦޻େɾҏ౻ڭतͱͷσΟεΧογϣϯ ҏ౻ѥࣿ ౦ژ޻ۀେֶ।ڭत ඒֶ w ࢹ֮ো֐ऀͱͷίϛϡχέʔγϣϯͷͨΊͷԠ༻ՄೳੑΛධՁ w ʮΠϧΧͷ஌֮ੈքʯΛମݧ͢Δ͜ͱͦͷ΋ͷ͚ͩͰ͸ͳ͘ɺ
 ҟͳΔ஌༷֮ࣜͷଘࡏΛର౳ͷ෺ͱͯ͠ೝΊΔ͜ͱʹ໘ന͕͋͞Δ

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4JHIU͸ ৽͍͠஌֮ੈք΁ͷύεϙʔτ = ʮ໨ʹΑΔݟ͑ʯΛ૬ରԽ͢Δ͜ͱͰ ҟͳΔ஌֮Λ࣋ͭਓʑͷؒͷཧղ΍ɺ ώτͷ஌֮ʹର͢ΔཧղΛਂΊΔ

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μΠΞϩάɾΠϯɾβɾμʔΫͱͷྨࣅ w ׬શͳ҉ҋͷதͰɺ
 ࢹ֮ো֐ऀͷΞςϯυʹಋ͔Ε
 ॳର໘ͷϝϯόʔͱڠྗ͠ͳ͕Β
 εϙʔπɾҿ৯ɾങ͍෺ͳͲΛମݧ w ීஈ໨Λ࢖͍ͬͯΔͱҙࣝͰ͖ͳ͍
 ࣗ෼ͷ஌֮ͷՄೳੑΛ஌Δ͜ͱ͕Ͱ͖Δ

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ಉ͡஌֮ੈքͷதͰ w 4JHIUΛ૷ணͨ͠··ɺεϙʔπ΍৯ࣄɺಃܳͳͲ
 ৽͍͠ମݧΛ௨ͨ͠ڞײ΍ڝ૪Λମݧ͢ΔϫʔΫγϣοϓΛ։͖͍ͨ

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͓ΘΓʹ

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·ͱΊ w ʮԻʹΑΔݟ͑ʯΛ࣮ݱͰ͖ΔσόΠεl4JHIUzΛ։ൃ w %γϛϡϨʔγϣϯʹΑΔ࿅श؀ڥ΋༻ҙ w Ҡಈɾ୳ࡧɾΩϟονϘʔϧͳͲͷλεΫΛ਱ߦ w ৽͍͠஌֮Λମݧ͢Δ͜ͱͰɺੜ·Ε͖ͭͷੈքͷݟ͑ํ Λ૬ରԽ͢ΔࢼΈl஌ཱྀ֮ߦzΛఏҊɾ࣮ݱ

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ँࣙ w ੴࠇߒ1. *1"ͷօ༷ w %..NBLF",*#" εϖʔεఏڙ w ౦ژେֶ($-ϦʔσΟϯάେֶӃ εϖʔεఏڙ w ౦ژܳज़େֶ৽Ҫᷞࢠಛ೚।ڭत w ౦ژ޻ۀେֶҏ౻ѥࣿڭत w ౦ژେֶኍ੉୩઒ݚڀࣨ ػࡐڠྗ w ϢʔβελσΟʹڠྗͯͩͬͨ͘͠͞ํʑ w Ճ౻༷ ࢁԼ༷ ௕ᖒ༷ w ౦ژେ੍ֶ࡞ల %$&910ͦͷ΄͔లࣔདྷ৔ऀͷΈͳ͞· w ը૾ΫϨδοτ w qJDLS.JLF"MMFO JDPOTDPN

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l஌ཱྀ֮ߦzͷͨΊͷࢹ֮୅ߦσόΠε ࿨Ոঘرླ໦ྑฏ ෬ݟྒྷฏफ૾༔ཬ "VEJUPSZWJTJPOTVCTUJUVUJPOEFWJDF GPSQFSDFQUJPOBMWPZBHF UIFTJHIUKQJOGP!UIFTJHIUKQ