0.3 0.2 0.1 0.0 0.1 0.2 0.3 0.4 PSE bias (µz,z⇤ ) Subject 1 0.4 0.5 0.6 0.7 0.8 0.9 1.0 1.1 0.15 0.10 0.05 0.00 0.05 0.10 0.15 Subject 2 v⇤ = 5, ⌧ = 100 v⇤ = 5, ⌧ = 200 v⇤ = 10, ⌧ = 100 v⇤ = 10, ⌧ = 200 1.0 1.1 1.2 1.3 0.4 0.5 0.6 0.7 0.8 0.9 1.0 1.1 1.2 1.3 0.15 0.10 0.05 0.00 0.05 0.10 0.15 Subject 2 v⇤ = 5, ⌧ = 100 v⇤ = 5, ⌧ = 200 v⇤ = 10, ⌧ = 100 v⇤ = 10, ⌧ = 200 Experimental Findings 307 308 309 310 311 312 313 314 315 316 317 318 319 320 321 322 323 is moving faster than a stimulus with parameters (v, z? ) . To asses the validity of our model, we tested four different scenarios by cons choices among z? = 0.78 c/ , v? 2 { 5 /s , 10 /s} , ⌧? 2 { 0.1s, 0.2s } , which corresponds to combinations of low/high speeds and temporal frequency b were generated on a Mac running OS 10.6.8 and displayed on a 20” Viewsonic resolution 1024 ⇥ 768 at 100 Hz. Routines were written using Matlab 7.10.0 and controlled the stimulus display. Observers sat 57 cm from the screen in a dark roo with normal or corrected to normal vision were used. They gave their inform experiments received ethical approval from the Aix-Marseille Ethics Committee the declaration of Helsinki. 3.2 Bayesian modeling To make full use of our MC paradigm in analyzing the obtained results, we follo of the Bayesian observer used for instance in [12]. We assume the observer mak a Maximum A Posteriori (MAP) estimator ˆ vz(m) = argmin v [ log( P M | V,Z(m | v, z)) log( P V | Z(v | z)) 6 281 282 283 284 285 286 287 288 289 290 291 292 293 294 295 296 297 298 299 300 301 302 303 304 305 3.1 Methods The task is to discriminate the speed v 2 R of MC stimuli moving w v0 = (v, 0) . We assign as independent experimental variable the aver we denote in the following z (we drop the index 0 to ease readabilit parameters are set to the following values V = 1 ⌧?z0 , ✓0 = ⇡ 2 , ⇥ = ⇡ 6 , Z = 0. Note that V is thus dependent of the value of z = z0 to ensure that ⌧? parameter ⌧? controls the temporal frequency bandwidth, as illustrated We used a two alternative forced choice (2AFC) paradigm. In each tri a small dark fixation spot was followed by two stimulus intervals of grey 250 ms inter-stimulus interval. The first stimulus has parameter has parameters (v2, z2) . At the end of the trial, a grey screen appears a which one of the two intervals was perceived as moving faster by pres is whether v1 > v2 or v2 > v1 . Given references values (v? , z? ) , for each trial, (v1, z1) and (v2, z2) ar ⇢ vi = v? , zi 2 z? + Z vj 2 v? + V , zj = z? where ⇢ V = { 2, 1, 0 Z = { 0.31, where (i, j) = (1, 2) or (i, j) = (2, 1) (i.e. the ordering is randomize values are expressed in cycles per degree (c/ ) and v values in /s. Ten possible combinations of these parameters are made per block of 250 blocks were collected per condition tested. The outcome of these exp psychometric curves ˆ 'v ? ,z ? , where for all (v v? , z z? ) 2 V ⇥ Z 283 284 285 286 287 288 289 290 291 292 293 294 295 296 297 298 299 300 301 302 303 304 305 306 307 The task is to discriminate the speed v 2 R of MC stimuli moving with a horizontal v0 = (v, 0) . We assign as independent experimental variable the average spatial freq we denote in the following z (we drop the index 0 to ease readability in this sectio parameters are set to the following values V = 1 ⌧?z0 , ✓0 = ⇡ 2 , ⇥ = ⇡ 6 , Z = 0.78 c/ . Note that V is thus dependent of the value of z = z0 to ensure that ⌧? = 1 V z0 stays parameter ⌧? controls the temporal frequency bandwidth, as illustrated on the left of F We used a two alternative forced choice (2AFC) paradigm. In each trial a grey fixatio a small dark fixation spot was followed by two stimulus intervals of 250 ms each, s grey 250 ms inter-stimulus interval. The first stimulus has parameters (v1, z1) and t has parameters (v2, z2) . At the end of the trial, a grey screen appears asking the partic which one of the two intervals was perceived as moving faster by pressing one of tw is whether v1 > v2 or v2 > v1 . Given references values (v? , z? ) , for each trial, (v1, z1) and (v2, z2) are selected so th ⇢ vi = v? , zi 2 z? + Z vj 2 v? + V , zj = z? where ⇢ V = { 2, 1, 0, 1, 2 } , Z = { 0.31, 0.15, 0, 0.16, 0. where (i, j) = (1, 2) or (i, j) = (2, 1) (i.e. the ordering is randomized across trials) values are expressed in cycles per degree (c/ ) and v values in /s. Ten repetitions of possible combinations of these parameters are made per block of 250 trials and at l blocks were collected per condition tested. The outcome of these experiments are s psychometric curves ˆ 'v ? ,z ? , where for all (v v? , z z? ) 2 V ⇥ Z , the value ˆ 'v ? empirical probability (averaged over the 40 trials) that a stimulus generated with para is moving faster than a stimulus with parameters (v, z? ) . 283 284 285 286 287 288 289 290 291 292 293 294 295 296 297 298 299 300 301 302 303 304 305 306 307 The task is to discriminate the speed v 2 R of MC stimuli moving with a horizo v0 = (v, 0) . We assign as independent experimental variable the average spatial f we denote in the following z (we drop the index 0 to ease readability in this se parameters are set to the following values V = 1 ⌧?z0 , ✓0 = ⇡ 2 , ⇥ = ⇡ 6 , Z = 0.78 c/ . Note that V is thus dependent of the value of z = z0 to ensure that ⌧? = 1 V z0 sta parameter ⌧? controls the temporal frequency bandwidth, as illustrated on the left We used a two alternative forced choice (2AFC) paradigm. In each trial a grey fix a small dark fixation spot was followed by two stimulus intervals of 250 ms eac grey 250 ms inter-stimulus interval. The first stimulus has parameters (v1, z1) an has parameters (v2, z2) . At the end of the trial, a grey screen appears asking the pa which one of the two intervals was perceived as moving faster by pressing one of is whether v1 > v2 or v2 > v1 . Given references values (v? , z? ) , for each trial, (v1, z1) and (v2, z2) are selected s ⇢ vi = v? , zi 2 z? + Z vj 2 v? + V , zj = z? where ⇢ V = { 2, 1, 0, 1, 2 } , Z = { 0.31, 0.15, 0, 0.16 where (i, j) = (1, 2) or (i, j) = (2, 1) (i.e. the ordering is randomized across tri values are expressed in cycles per degree (c/ ) and v values in /s. Ten repetitions possible combinations of these parameters are made per block of 250 trials and blocks were collected per condition tested. The outcome of these experiments ar psychometric curves ˆ 'v ? ,z ? , where for all (v v? , z z? ) 2 V ⇥ Z , the value ' empirical probability (averaged over the 40 trials) that a stimulus generated with p is moving faster than a stimulus with parameters (v, z? ) . 283 284 285 286 287 288 289 290 291 292 293 294 295 296 297 298 299 300 301 302 303 304 305 306 307 The task is to discriminate the speed v 2 R of MC stimuli moving with a hori v0 = (v, 0) . We assign as independent experimental variable the average spatia we denote in the following z (we drop the index 0 to ease readability in this parameters are set to the following values V = 1 ⌧?z0 , ✓0 = ⇡ 2 , ⇥ = ⇡ 6 , Z = 0.78 c/ . Note that V is thus dependent of the value of z = z0 to ensure that ⌧? = 1 V z0 parameter ⌧? controls the temporal frequency bandwidth, as illustrated on the le We used a two alternative forced choice (2AFC) paradigm. In each trial a grey a small dark fixation spot was followed by two stimulus intervals of 250 ms e grey 250 ms inter-stimulus interval. The first stimulus has parameters (v1, z1) has parameters (v2, z2) . At the end of the trial, a grey screen appears asking the which one of the two intervals was perceived as moving faster by pressing one is whether v1 > v2 or v2 > v1 . Given references values (v? , z? ) , for each trial, (v1, z1) and (v2, z2) are selected ⇢ vi = v? , zi 2 z? + Z vj 2 v? + V , zj = z? where ⇢ V = { 2, 1, 0, 1, 2 } , Z = { 0.31, 0.15, 0, 0 where (i, j) = (1, 2) or (i, j) = (2, 1) (i.e. the ordering is randomized across values are expressed in cycles per degree (c/ ) and v values in /s. Ten repetitio possible combinations of these parameters are made per block of 250 trials an blocks were collected per condition tested. The outcome of these experiments psychometric curves ˆ 'v ? ,z ? , where for all (v v? , z z? ) 2 V ⇥ Z , the valu empirical probability (averaged over the 40 trials) that a stimulus generated wit is moving faster than a stimulus with parameters (v, z? ) . P Z(z) / z e , P ⇥(✓) / e and P || V v0 ||(r) / e . (6) z0 Z V ⇠1 ⌧ ⇠2 ⇠1 ✓0 z0 ⇥ Z Slope: \ v0 Two different projections of ˆ in Fourier space t MC of two different spatial frequencies z Figure 2: Graphical representation of the covariance (left) —note the cone-like shape of the envelopes– and an example of synthesized dynamics for narrow-band and broad-band Motion Clouds (right). Plugging these expressions (6) into the definition (5) of the power spectrum of the motion cloud, one obtains a parameterization which is very similar to the one originally introduced in [9]. The fol- lowing table articulates the speed v0 and frequency (✓0, z0) central parameters in term of amplitude and orientation, each one being coupled with the relevant dispersion parameters. Figure 2 shows a graphical display of the influence of these parameters. Speed Freq. orient. Freq. amplitude (mean, dispersion) (v0, V ) (✓0, ⇥) (z0, Z) Remark 2. Note that the final envelope of ˆ is in agreement with the formulation that is used in [8]. However, this latter derivation was based on a heuristic following a trial-and-error strategy between modelers and psychophysicists. Herein, we justified these different points in a constructive manner. P Z(z) / z e , P ⇥(✓) / e and P || V v0 ||(r) / e . (6) z0 Z V ⇠1 ⌧ ⇠2 ⇠1 ✓0 z0 ⇥ Z Slope: \ v0 Two different projections of ˆ in Fourier space t MC of two different spatial frequencies z Figure 2: Graphical representation of the covariance (left) —note the cone-like shape of the envelopes– and an example of synthesized dynamics for narrow-band and broad-band Motion Clouds (right). Plugging these expressions (6) into the definition (5) of the power spectrum of the motion cloud, one obtains a parameterization which is very similar to the one originally introduced in [9]. The fol- lowing table articulates the speed v0 and frequency (✓0, z0) central parameters in term of amplitude and orientation, each one being coupled with the relevant dispersion parameters. Figure 2 shows a graphical display of the influence of these parameters. Speed Freq. orient. Freq. amplitude (mean, dispersion) (v0, V ) (✓0, ⇥) (z0, Z) Remark 2. Note that the final envelope of ˆ is in agreement with the formulation that is used in [8]. However, this latter derivation was based on a heuristic following a trial-and-error strategy between modelers and psychophysicists. Herein, we justified these different points in a constructive manner. µz,z? z z µz,z?